2007). The vast majority of trendy domesticated rice varieties lack proanthocyanidin pigments within the pericarp; this domestication trait arose through human selection for loss-of-function mutations on the Rc locus, which encodes a bHLH regulatory protein in the proanthocyanidin synthesis pathway (Sweeney et al. 2006). Selection for non-pigmented grains within the crop might probably reflect human aesthetic preferences, selection against seed dormancy, selection for improved style and cooking qualities, and/or choice for different domestication traits linked to Rc (Gu et al. 2004; Sweeney et al. 97% of non-pigmented cultivars (Sweeney et al. 2007), is characterized by a 14-bp frameshift deletion in exon 7 (initially annotated as exon 6; see Sweeney et al. 2006; Furukawa et al. 2007). This mutation generates a truncated, nonfunctional gene product and the nonpigmented (‘white’) pericarp of domesticated rice (Sweeney et al. 2006; Furukawa et al. 2007). An independently advanced domestication allele, Rc-s, is characterized by a C→A substitution in exon 7 that creates a premature cease codon (Sweeney et al.
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